that's a tougher one.
The Acela Express isn't a good example, because (I believe) the cars are coupled together with something more akin to a drawbar (no slack action) or an MU coupler than a conventional knuckle coupler. I'm not sure on that, but there is essentially no slack action in them that you can feel as a passenger. I'll have to hope that someone with more shop-wise knowledge of the AE's can answer that.
Then, when there are units on both end of a train, the can only both be working if the coaches are equipped for push-pull service. If so, the engines can be MUed through the coaches (just as a cab car is essentially "MUed" to its pushing locomotive through the coaches). However, if two units are on each end of the train, and the coaches are not push-pull equipped, then the trailing engine is simply dead in tow (either shut down, or idling, but isolated, and just along for the ride).
That's the layman's explanation. As for the details of how slack action is controlled, I would imagine that its a similiar challenge to managing the slack when pushing with a push pull. Only a passenger enginner can really explain that.
Wondering if I'll see the Western Route double-tracking finished before I retire...
Photo: Melbourne W7 No. 1019 on Route 78, Bridge & Church Streets, Richmond, Victoria. 10/21/2010